| 
            Hansen EC. Recherches sur la physiologie et la morphologie des ferments alcooliques. XIII. Nouvelles etudes sur les levures de brasserie a fermentation basse.  C.R. Trav. Lab. Carlsberg 7: 179-217, 1908. 
         
            Piskur J, et al. A mitochondrial molecular marker, ori-rep-tra, for differentiation of yeast species.  Appl. Environ. Microbiol. 61: 2780-2782, 1995. PubMed: 7618892
         
            Walther A, Hesselbart A, Wendland J. Genome Sequence of Saccharomyces carlsbergensis, the World's First Pure Culture Lager Yeast. G3 (Bethesda) 4: 783-793, 2014. PubMed: 24578374
         
            Huang CH, Lee FL, Tai CJ. The beta-tubulin gene as a molecular phylogenetic marker for classification and discrimination of the Saccharomyces sensu stricto complex. Antonie van Leeuwenhoek 95: 135-142, 2009. PubMed: 19112604
         
            Ogata T, et al. Chromosomal location of Lg-FLO1 in bottom-fermenting yeast and the FLO5 locus of industrial yeast. J. Appl. Microbiol. 105: 1186-1198, 2008. PubMed: 18564345
         
            Nguyen HV, Gaillardin C. Evolutionary relationships between the former species Saccharomyces uvarum and the hybrids Saccharomyces bayanus and Saccharomyces pastorianus; reinstatement of Saccharomyces uvarum (Beijerinck) as a distinct species. FEMS Yeast Res. 5: 471-483, 2005. PubMed: 15691752
         
            Salema-Oom M, et al. Maltotriose utilization by industrial Saccharomyces strains: characterization of a new member of the alpha-glucoside transporter family. Appl. Environ. Microbiol. 71: 5044-5049, 2005. PubMed: 16151085
         
            Kurtzman CP, Robnett CJ. Phylogenetic relationships among yeasts of the 'Saccharomyces complex' determined from multigene sequence analyses. FEMS Yeast Res 3: 417-432, 2003. PubMed: 12748053
         
            Okuda Y, et al. Occurrence, horizontal transfer and degeneration of VDE intein family in Saccharomycete yeasts. Yeast 20: 563-573, 2003. PubMed: 12734795
         
            Perez-Ortin JE, et al. Molecular characterization of a chromosomal rearrangement involved in the adaptive evolution of yeast strains. Genome Res. 12: 1533-1539, 2002. PubMed: 12368245
         
            Casaregola S, et al. Analysis of the constitution of the beer yeast genome by PCR, sequencing and subtelomeric sequence hybridization. Int. J. Sys.t Evol. Microbiol. 51: 1607-1618, 2001. PubMed: 11491364
         
            Goncalves PM, et al. FSY1, a novel gene encoding a specific fructose/H+ symporter in the type strain of Saccharomyces carlsbergensis. J. Bacteriol. 19: 5628-5630, 2000. PubMed: 10986274
         
            type strain of Saccharomyces carlsbergensis
         |